Sqn. Ldr. Brian Lane was the Officer Commanding No.19 Squadron, based at Fowlmere in September 1940 and was flying this aircraft when he shot down a Messerschmitt Bf.110 of Stab.II./ZG 2 near North Weald on 7th September. Two days later, the same aircraft shot down another Bf.110, whilst being flown by Flt. Lt. Peter Lawson. On September 11th it also shot own a Heinkel He-111. After a brief period with No.152 Sqn., the Spitfire went to No. 58 OTU in 1941, and continued to perform second-line tasks until it was written-off in May 1944 whilst serving with No. 57 OTU. The aircraft wore a non-standard yellow prop spinner reputedly painted during its brief OTU service.